The "local patch quality perspective" revealed the importance to C. rufocanus of a large patch of forest >60 yr old containing a lot of old-growth (pine) forest (>100 yr). In fact, at the landscape level, the frequency distribution of focal patches of forest >60 yr old and especially their content of old-growth pine forest (>100 yr), relative to the properties of plots with C. rufocanus, suggests that there are few forest patches left that are suitable for C. rufocanus in our study area. Our results strongly suggest habitat fragmentation as a contributing cause to the long-term decline of C. rufocanus.